It is the year 1214. A baby squirrel falls from the nest and lands in the passing saddlebag of a 13-year-old boy. Still blind deaf hairless and unable to eat solid food there is little hope for Puttermunch's survival. Moved by the persistent pleading of his distraught mother a selection of five squirrels set out to find Puttermunch launching a perilous and life-altering journey on the Way of St. James in Spain. The road is fraught with failure for Billowtail Tippy Nip Sugarcoat and Sir Sniff until these well-meaning tree squirrels born to be loners and hoarders learn the value of community and self-sacrifice.